Gould, The Ediacaran Experiment

the Ediacaran animals with modern groups. Thus, we have Ediacaran jellyfish, corals, and worms -a continuity of evolutionary relationship across the greatest of all geological boundaries. Yet, as I argued just a few months ago in my column on conodonts (July 1983), the traditional ploy of forcing old and problematical animal fossils into modern taxonomic categories often fails badly. We must recognize that the early history of life should be studded with failed experiments--small groups that never achieved much diversity and bear only distant relationship with any modern animal. We might expect that our oldest fauna should contain a large number of such curiosities--yet all Ediacaran animals have been shoehorned, often with considerable effort, into modern groups.

Dolf Seilacher now argues, turning the old view completely on its head, that the Ediacaran fauna contains not simply a few creatures with no modern analogues--but that every animal in it shares a basic mode of organization quite distinct from the architecture of living groups. The entire Ediacaran fauna, in other words, represents a unique and extinct experiment in the basic construction of living things. Our planet's first fauna was replaced after a mass extinction, not simply improved and expanded.

Dolf began by showing that the similarities of Ediacaran and modern animals are misleading and superficial, and that the Ediacaran forms could not work as their supposed living counterparts. Nearly all Ediacaran fossils have been falsely fit into three modern groups: jellyfish, corals, and segmented worms. Living jellyfish move by contracting a prominent ring of concentric muscles located at the outer edge of their bell, radial grooves for feeding lie within the concentric muscles, toward the center. But the so-called Ediacaran medusoids have a reversed arrangement that could not work in the same way: concentric structures surround the center, and radial grooves lie on the outside.

Modern alcyonarian corals ("soft" corals, or sea pens) invariably bear distinct branches, often springing from a common stem. The branches must be separated so that water, bearing oxygen and nutrients, can reach the individual polyps (members of the colony) growing on them. At first glance, the Ediacaran "sea pens" look superficially like their modern counterparts in general shape, but they form a continuous, quilted structure, not a set of separated branches--and could therefore not operate like a modern soft coral colony. The Ediacaran "worms" are segmented and bilaterally symmetrical like their supposed modern analogues, but so are many other creatures--and such a basic and repeatable architecture need not imply close relationship. In other respects, the Ediacaran creatures are most unwormlike. They may be up to a meter in length and flat as a pancake--more like films than the substantially thickened bodies of most modern segmented worms.

After tracing the differences between Ediacaran animals and their supposed modern counterparts, Seilacher examined the similarities that unite all Ediacaran forms. They seem to share an architecture only rarely utilized by modern animals--and not by any living creature ever linked to an Ediacaran fossil. They look like ribbons, pancakes, and films, sometimes slightly "blown up" as air mattresses with a foliate or quilted structure.

The Ediacaran animals evolved before any creature had invented mineralized skeletons or any external hard parts. Perhaps their unique Bauplan (to use the convenient German term for a basic scheme of organic architecture), or "building plan," records a pathway to large size that animals without supporting hard parts might follow--light and thin structures, woven together for added strength. In any case, and following a favorite theme of these columns for more than a decade, the Ediacaran fossils seem to represent one of two possible solutions--the one not followed by modern animals--to the basic structural problem of large size: the imposed decline of surfaces relative to volumes since surfaces (growing as length squared) must increase more slowly than volumes (growing as length cubed) as objects of similar shape get bigger. Since so many organic functions depend upon surfaces (respiration and feeding, to name just two) yet must serve the entire body's volume, this decline in relative surface cannot be tolerated for long.

Of the two possible solutions, most modern animals have retained their rounded or globular shapes but have evolved internal organs to increase surface areas--lungs for respiration and the complexly folded surface of the small intestine for absorption of food, for example. Another potential solution, followed rarely today but exploited by some large parasites, including tapeworms, permits large size without any internal complexity by changing the body's basic shape into something very thin--a ribbon or pancake--so that no internal space will be far from the external surface, the only locus of respiration and absorption of food in the absence of internal organs. The Ediacaran animals, as a group, have followed this second pathway to large size and therefore

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